In 1961, the Ark, a Cybertronian spacecraft carrying an invention capable of ending the war between the Autobots and the Decepticons, crash lands on the dark side of Earth's Moon. The crash is detected on Earth by NASA, and President John F. Kennedy authorizes a mission to put a man on the Moon as a cover for investigating the spacecraft. In 1969, the crew of Apollo 11 lands on the Moon and secretly inspect the Ark before heading home. President Richard Nixon orders them to keep the details of this mission secret.
In the present day, four years
after Egypt [N 1], the Autobots assist the humans in
preventing major conflicts. During a mission to the site of the 1986
Chernobyl nuclear disaster in Ukraine to investigate suspected
alien technology, the Autobots are attacked by Decepticon scientist Shockwave and
his giant worm Driller. After the two escape, Optimus Prime discovers
that the technology is a fuel cell from the Ark, thought to have
been lost after escaping Cybertron. After learning of the top-secret mission to
the Moon, the Autobots travel there and discover Sentinel Prime—Optimus's
predecessor as leader of the Autobots—in a comatose state, as well as five
Pillars he created as a means of establishing a "Space Bridge", which
is a wormhole that can be used to teleport matter between two points.
Upon returning to Earth, Optimus uses the energy of the Matrix of
Leadership to revive Sentinel Prime.
Meanwhile, Sam Witwicky lives
with his new girlfriend, Carly Spencer but is unable to work with the
Autobots or find a job. After Sam finds a job, his co-worker, Jerry Wang
provides him information about the Ark, before Jerry is
assassinated by the Decepticon Laserbeak. Sam contacts Seymour Simmons, and
they realize the Decepticons are murdering people connected to the American and
Soviet space missions to the Ark. They locate two surviving
cosmonauts, who reveal satellite photos of hundreds of Pillars being stockpiled
on the Moon. Sam realizes that the Decepticons raided the Ark long
before the Apollo 11 mission and intentionally left Sentinel and the five
Pillars for the Autobots to find, as they know that Sentinel is the key to
activating the Pillars. The Autobots fight off the pursuing Decepticons and
rush to return Sentinel to their base for protection, but Sentinel betrays them
and kills Ironhide, revealing that he had made a deal with Megatron to
ensure Cybertron's survival.
Sentinel uses the Pillars to
transport hundreds of concealed Decepticons from the Moon to Earth. Dylan
Gould, Carly's boss, is revealed to be working with the Decepticons after he
captures Carly with the help of Soundwave, who had been masquerading as
a Mercedes SLS AMG he gifted to Carly. The Autobots are exiled from
Earth at the demand of the Decepticons, but as their ship leaves Earth, it is
destroyed by Starscream, seemingly killing the Autobots. The Decepticons
invade Chicago as they place Pillars around the world to begin
transporting their homeworld Cybertron to the Solar System, and then use
Earth's resources to rebuild their world, including humans, who will used
as slave labor. Sam teams up with former soldier Robert Epps to go into
Chicago to save Carly and arrest Dylan, but they are nearly killed by the
Decepticons before the Autobots intervene, revealing that the Autobots
concealed themselves during the launch of their ship to convince the
Decepticons they were destroyed.
The Autobots, along with NEST
teams and Navy SEALs, manage to rescue Carly and begin fighting off the
Decepticons. Soundwave and the Decepticons temporarily incapacitate and capture
a few Autobots, killing Que. However, the miniature Autobots Wheelie and Brains
sabotage a Decepticon ship, creating a distraction and allowing the Autobots to
escape, with Bumblebee killing Soundwave in the process. Many other
Decepticons, including Laserbeak, the Driller, Starscream, and Shockwave, are
killed in the battle.
Sam fights Dylan and knocks him
into the Pillar, electrocuting and killing him. Bumblebee and Ratchet
destroy the Control Pillar, permanently disabling the Bridge and causing the
partially transported Cybertron to be destroyed after being imploded. Optimus
and Sentinel fight while Carly convinces Megatron that he will be replaced by
Sentinel as the leader of the Decepticons. Sentinel gains the upper hand
against Optimus but is incapacitated by Megatron, who is determined to regain
his leadership. Megatron then falsely offers a truce, but Optimus refuses to
accept his surrender and decapitates him. An indifferent Optimus then executes
Sentinel, despite his pleas for mercy. Carly and Sam are reunited, and the
Autobots remain on Earth as their permanent home.
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