Thousands of years ago, the planet Cybertron was consumed by a civil war between the two Transformer factions, the Autobots led by Optimus Prime and the Decepticons led by Megatron. The Autobots want to find the All Spark, the source of all Cybertronian life, so they can use it to rebuild Cybertron and end the war between the Autobots and the Decepticons, while the Decepticons want to use it to defeat the Autobots and conquer the universe. Megatron found the All Spark on Earth, but crash-landed in the Arctic Circle and was frozen in the ice. Captain Archibald Witwicky and his crew of explorers stumble upon Megatron's body in 1897. Captain Witwicky accidentally activates Megatron's navigational system, causing his eyeglasses to be imprinted with the coordinates of the All Spark's location. Sector 7, a secret United States government organization, discovers the All Spark in the Colorado river and builds the Hoover Dam around it to mask its energy emissions. The still-frozen Megatron is moved into this facility and is reverse engineered to advance human technology.
In the present day, the
Decepticons—Blackout, Scorponok, Frenzy, Barricade, Starscream, Brawl and Bonecrusher—have
landed on Earth and assumed the disguise of Earth vehicles. Blackout and
Scorponok attack the U.S. SOCCENT military base in Qatar and
try to hack into the U.S. military network to find the location of
Megatron and the All Spark. Their mission is thwarted when the base staff severs
the network cable connections. While Blackout destroys the rest of the base,
Scorponok chases a small group of survivors who have photographic evidence of
the robots, but he is eventually repelled. During this battle, the military
discovers its only effective weapons against the Transformers' armor are
high-heat sabot rounds.
After Blackout's failure, Frenzy
infiltrates Air Force One to try again to hack into the military
network, and in doing so plants a virus. He finds the map imprinted on Captain
Witwicky's glasses, whose descendant, Sam Witwicky, intends to sell on eBay.
Frenzy and Barricade begin tracking Sam's location. The Autonomous Robotic
Organism (shortened to "Autobot") Bumblebee is also on
Earth, disguised as a 1976 Chevrolet Camaro,[6] and
is bought by Sam while shopping for his first car. Bumblebee helps him woo his
crush, Mikaela Banes. Bumblebee leaves at night to transmit a homing
signal to the rest of the Autobots and Sam sees him in robot mode. Barricade
confronts Sam and demands Archibald's spectacles, but Bumblebee rescues him and
Mikaela. After Bumblebee upgrades his vehicle form by scanning a 2006 Chevrolet
Camaro, they leave to rendezvous with the rest of the Autobots: Optimus
Prime, Jazz, Ironhide, and Ratchet—who have landed on Earth and
taken the forms of Earth vehicles as well. Sam, Mikaela, and the Autobots
return to Sam's home and obtain the glasses. Agents from Sector 7 arrive and
capture Sam, Mikaela, and Bumblebee.
Frenzy, disguised as a cellphone,
secretly accompanies the group to Hoover Dam and releases Megatron from his
frozen state. Locating the All Spark, Frenzy sends an alert to the other
Decepticons. Sam convinces the Sector 7 agents to release Bumblebee so that he
can get the All Spark to Optimus Prime. Frenzy's virus has shut down government
communications, but a pair of hackers manage to establish a signal to the Air
Force. The Autobot-human convoy goes to nearby Mission City to obtain a radio
that will guide the Air Force's defense and secure a rendezvous point as a safe
destination for All Spark. The Decepticons attack and Bonecrusher, Frenzy,
Jazz, Devastator, and Blackout are all killed during the ensuing battle, but
Sam manages to ram the All Spark into Megatron's chest, killing Megatron and
destroying the All Spark. Optimus takes a fragment of the All Spark from
Megatron's corpse but realizes that with its destruction, their homeworld
Cybertron cannot be restored. Consequently, Optimus sends a signal to other
surviving Autobots in the universe, directing them to their new home, Earth.
The government orders the decommission of Sector 7 and has the deceased
Decepticons dumped into the Laurentian Abyss. In the post-credits,
Starscream, who fled the battle, escapes into space.



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