In 17,000 B.C., the Primes (the original Autobots) got their energy from sun harvesters, machines that destroy stars to harness their energy...

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Transformer: The Revenge of the Fallen (2009)

In 17,000 B.C., the Primes (the original Autobots) got their energy from sun harvesters, machines that destroy stars to harness their energy. The Primes vowed never to destroy a star that sustains life. One Prime attempted to violate this rule by building a sun harvester on Earth, for which he was imprisoned by the other six Primes, becoming "The Fallen", the original Decepticon.

In 2009, two years after the battle of Mission City,[N 1] the Autobots and the humans have formed the Non-biological Extraterrestrial Species Treaty (NEST), a classified international task force meant to eliminate the remaining Decepticons who are hunting the Autobots. They defeat two Decepticons, Sideways and Demolisher, in Shanghai, but the latter tells them the Fallen is coming before being killed. Meanwhile, the Decepticon Soundwave hacks into a military satellite. The Decepticons steal the last known piece of the AllSpark shard from a Navy base and use it to resurrect Megatron's body while tearing off parts from one of their own members to provide necessary repairs to his body. The Fallen sends Megatron and his second-in-command, Starscream, to capture Sam Witwicky alive and kill Optimus.

Sam, now a college student, has been seeing Cybertronian symbols since holding a smaller AllSpark shard; Megatron believes the symbols will lead the Decepticons to a new Energon source. The shard brings many of the kitchen appliances to life and they try to kill Sam and his parents; Bumblebee rescues them. Sam gives the shard to his girlfriend Mikaela Banes, who later captures the Decepticon Wheelie as he attempts to steal it. After being attacked by Alice, a Decepticon Pretender posing as a college student, Sam, his roommate Leo, and Mikaela are captured by the Decepticon Grindor before Optimus and Bumblebee rescue them. The Decepticons then kill Optimus (but not before he kills Grindor) and launch devastating, simultaneous attacks around the world, while the Fallen, with help from Soundwave, hijacks Earth's telecommunications systems, demanding that Sam be handed over to him.

Sam, Mikaela, and Leo then find alien expert and former Sector Seven agent, "Robo-Warrior" Seymour Simmons, who reveals the Transformers visited Earth eons ago and the most ancient, known as Seekers, remained hidden on Earth. With help from Wheelie, they track down an elderly Decepticon turned Autobot Seeker named Jetfire at the Smithsonian Air and Space Museum. They use their shard to revive Jetfire, who teleports the group to Egypt. Along with Jetfire, Wheelie sides with the Autobots, and Jetfire sends them to locate the Matrix of Leadership, the sun harvester's key, which could also be used to revive Optimus. The group finds the Matrix, whom the Primes that punished the Fallen sacrificed themselves to hide, in Petra, but it disintegrates into dust.

Meanwhile, NEST forces and the Autobots land near the Giza pyramid complex and are attacked by the Decepticons. The Constructicons combine to form Devastator, who destroys one of the pyramids to reveal the sun harvester inside before he is killed by the navy. A majority of the Decepticons are killed, mostly with multiple airstrike from the Navy and the Air Force, but Megatron kills Sam. Near-death, the Primes speak to Sam, saying that the Matrix must be earned, not found and that he has now earned the right to bear it by fighting for Optimus. They revive Sam and grant him the Matrix, which he uses to revive Optimus. The Fallen steals the Matrix from a weak Optimus and uses it to activate the sun harvester. After Jetfire (who was wounded by Scorponok, but was able to kill his attacker) sacrifices himself to allow his parts to be transplanted to Optimus, Optimus Prime destroys the harvester and kills the Fallen; Megatron and Starscream retreat. The Autobots and their allies then return to the United States, and Leo and Sam return to college.

 

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