In 2015, aliens called "Mimics" arrive in Germany via an asteroid and swiftly conquer most of continental Europe. By 2020, the United Defense Force (UDF), a global military alliance established to combat the alien threat, finally achieves a victory over the Mimics at Verdun using newly developed mech-suits. In Britain, the UDF plans a major invasion of France, and General Brigham orders recently attached U.S. Army Major William Cage, a public affairs officer, to cover it. Cage, having no combat experience, objects and threatens to blame Brigham if the invasion fails. Brigham has Cage arrested and sent to Heathrow Airport, now a military base. Cage awakens to find Brigham has demoted him to a private and assigned him to Master Sergeant Farell and the misfit J-Squad, who do not take kindly to Cage.
The landing in France is a
colossal failure; Farell and J-Squad are quickly killed. Cage uses a Claymore
mine to kill an unusually large blue Mimic but is mortally wounded by the
explosion and covered in the alien's blood. Cage jolts awake to find himself
back at Heathrow, reliving the events of the previous morning. His attempts to
warn Farell against the invasion are ignored, and he experiences the loop of
dying on the beach and waking up at Heathrow repeatedly. During one loop, Cage
tries to save Sergeant Rita Vrataski, a celebrated hero of the battle of
Verdun. She realizes Cage can loop time and orders him to find her the next
time he wakes up.
Cage reawakens and locates
Vrataski, who takes him to Dr. Carter, an expert in Mimic biology. He explains
that the Mimics are a superorganism in which the "Omega"
controls the cerebrum, while the "Alphas" behave as the ganglia through
which the Omega controls ordinary Mimics; if an Alpha is terminated, the Omega
resets the day and adjusts its tactics until the battle is won. Cage
inadvertently "hijacked" their ability to reset time through his
exposure to an Alpha's blood. Vrataski had this ability at Verdun, using it to
win the battle before she was wounded and received a blood transfusion,
losing the power. She tells Cage to locate and kill the Omega to end the alien
invasion.
Over many more loops, Vrataski
trains Cage to excel in combat. After a frustrating lesson, Cage escapes to
London, only to discover that the Mimics will attack there next after the
invasion. After seeing visions of a dam in Germany where the Omega is hiding
and spending many loops figuring out how to escape the invasion and reach the
dam, Cage grows closer to Vrataski, but she is only interested in the mission.
Cage flies to the dam alone upon reaching a point where Vrataski is killed no
matter what they do. The Omega is not there and he is ambushed by an Alpha who
attempts to strip him of his ability to reset time, but Cage deliberately
drowns himself.
Cage and Vrataski infiltrate
the Ministry of Defence, where Cage convinces Brigham to give him Carter's
prototype device that can locate the Omega, but they are pursued by military
police on leaving. During the ensuing car chase, Cage uses the device and
discovers the Omega is under the Louvre Pyramid in Paris. Cage is
seriously injured during capture and wakes up in a hospital to find he has been
given a blood transfusion and has lost the ability to loop time.
Vrataski frees Cage and they
recruit J-Squad to help destroy the Omega before the invasion begins. They fly
to Paris, where the soldiers sacrifice themselves so that Cage and Vrataski can
reach the Louvre. Before luring away an Alpha standing between them and the
submerged Omega, Vrataski kisses Cage to thank him for getting her as far as he
did. When the Alpha kills Vrataski and mortally wounds Cage, he manages to drop
a belt of grenades that destroys the Omega, killing the Alpha and the Mimics.



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