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Brothers Sam and Nathan "Nate" Drake are caught by museum security trying to steal the first map made after the Magellan expedition. The orphanage that houses both boys kicks Sam out. Before he leaves, Sam promises his return to Nate, and leaves him a ring with the inscription "Sic Parvis Magna".
Fifteen years later, Nate works as a bartender in New
York City and pickpockets wealthy patrons. Victor "Sully"
Sullivan, a fortune hunter who worked with Sam tracking treasure hidden by the
Magellan crew, explains to Nate that Sam vanished after helping him steal Juan
Sebastian Elcano's diary. Nate, who received several postcards from Sam over
the years, agrees to help Sully to find his brother. Sully and Nathan go to an
auction to steal a golden cross linked to the Magellan crew, where they meet
Santiago Moncada, the last descendant of the Moncada family who funded
Magellan's expedition, and Jo Braddock, leader of mercenaries hired by Moncada.
Nate is ambushed by Braddock's men, and the ensuing fight creates a distraction
for Sully to steal the cross.
The duo travel to Barcelona, where the treasure is
supposedly hidden, and rendezvous with Sully's contact Chloe Frazer, who
has another cross. Chloe steals the first cross from Nate, but the men convince
her to work with them. Meanwhile, Moncada learns from his father that the
family fortune will be donated, and so orders Braddock to kill his father, so
that he will inherit the money instead. Nate, Chloe, and Sully follow clues in
Elcano's diary to Santa Maria del Pi, finding a secret crypt behind the
altar. Nate and Chloe enter, finding a trap door, but as they open it, the
crypt floods with water. Sully helps them escape after subduing an ambush by
Braddock. Using the two crosses to unlock a secret passage, Nate and Chloe find
a map that indicates the treasure is in the Philippines. Chloe betrays
Nate and takes the map, hinting Sully is keeping a secret about Sam. Chloe
brings the map to Moncada, having also been hired by him, to Braddock's
displeasure.
Sully recovers Nate and says after he and Sam recovered Elcano's
diary, they were ambushed by Braddock; Sam was shot and Sully narrowly escaped.
Moncada, Chloe, and Braddock's team depart in a cargo plane to find the
treasure, but Braddock kills Moncada to gain power, forcing
Chloe into hiding with the map. Nate and Sully secretly board the plane and
Nate confronts Braddock with Chloe's help while Sully parachutes out. Nate and
Chloe are ejected from the plane during the ensuing battle and the pair land in
the Philippines, where they realize the map does not pinpoint the treasure.
Nate discovers the treasure's true location through hints left by Sam's
postcards, but leaves fake coordinates for Chloe after correctly doubting her
loyalties. He discovers the Magellan ships and reunites with Sully. Braddock
follows them, forcing Nate and Sully to hide as her crew airlift the ships.
In their escape, Sully commandeers one of the helicopters,
causing Braddock to order the other helicopter to approach for a boarding
action. Nate defends himself from her mercenaries and shoots down the other
helicopter with one of the ship's cannons. Braddock drops that ship's anchor
while Nate climbs to the helicopter. Sully throws a bag of collected treasure
at Braddock who falls into the sea and is crushed to death when the ship breaks
and falls on her. As Philippine naval units arrive, Nate and Sully
get away with a few pieces of pickpocketed treasure while Chloe is left empty
handed as she decides to follow the duo. It is then revealed that in a Prison
somewhere, Nate's brother Sam is seen writing a postcard, warning his brother
to be careful of those around him. A second scene shows Nate meeting with Gabe
in a bar, someone who works for Roman. Sully comes in to save Nate after being
held up by Gabe, and was caught again after leaving the bar, possibly leading
to a potential sequel.


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